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The Influences of Queen Mandibular Pheromone and Brood Pheromones on the Process of Queen Rearing in Colonies of Honey Bees The Influences of Queen Mandibular Pheromone and Brood Pheromones on the Process of Queen Rearing in colonies of Honey Bees (Apis meliferra spp.)Highly social colonies of honeybees (Apis mellifera spp.) communicate through the use of.