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How Does the Reducing Equivalents Go from the Cytoplasm to the Mitochondrion? How does the reducing equivalents go from the cytoplasm to the mitochondrion? They don’t cross the membrane, but electrons do through two ways:Glycerol 3-phosphate shuttle:In cytoplasm, DHAP is reduced to glycerol 3-phospahe by NADH. Enzyme: cytoplasmic Glycerol 3-phoshate dehydrogenase.FAD is reduced to FADH2 in.